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find no characters worthy to recognize two species here. Based on the description and type specimen photo, I likewise reduce the Chinese species, which falls within the range of variation for this species.

E. Merremia thorelii (Gagnep.) Staples, comb. nov.Ipomoea thorelii Gagnep., Notul. Syst. (Paris) 3: 1915. Types: Vietnam. Ti-Tinh, Thorel s.n. (syntype, P!, 2 sheets). Laos. Khong, Thorel s.n. (syntype, P!).— Merremia collina S.Y.Liu, Guihaia 4: 199. 1984, synon. nov. Type: China, Guangxi,Yongning Xian, 5 Oct. 1983, S.Y. Liu 002112 (holotype, GXCM n.v.; isotypes IBK n.v., KUN n.v.).

The protologue and original illustration leave no doubt that the Chinese plant is conspecific with the Indochinese species first named as I. thorelii.


Two species identified by Traiperm in her unpublished M.Sc. thesis (Traiperm 2002) have been rigorously checked by Staples and proven to be undescribed. They are described and given scientific names here.

A.Argyreia variabilis Traiperm & Staples, sp. nov., simillima A. kerrii sed lamina angustior, basin non cordatis, duplo facies confertim brunneopilosis institiae, et sepalis latior obovatis orbicularis, totius glabris differt. Type: Thailand. Phetchaburi: Kaeng Krachan National Park, 1 Sept. 2001, Traiperm 31 (holotype BKF; isotypes—to be distributed—A, BCU, K, KKU). Fig. 4 & Fig. 6.

Woody climber, stems to 30 m tall, terete, densely brown pubescent to hirsute, internodes 1–11 cm long. Leaves ovate or ovate-lanceolate, 10–24 by 4–10 cm, base obtuse or rounded, rarely truncate, margins entire, apex acute to acuminate, chartaceous; upper surface densely brown hirsute; lower surface densely brown pilose; lateral nerves 14–16 pairs, prominent beneath; petiole 1.5–5.0 cm long, grooved, densely brown-pubescent. Inflorescences axillary, lax, several-flowered cymes; peduncles 2–3.5 cm long, terete, densely brown-strigose; pedicels 5–10 mm long, densely brown-strigose; bracts lanceolate to elliptic- lanceolate, 15–25 by 5–8 mm, apex acute to acuminate, densely brown pubescent outside, glabrous inside, persistent. Flower sepals subequal, entire, broadly obovate to orbicular, 15–17 by 11–13 mm, glabrous on both sides; corolla fleshy, campanulate, 6–7 cm long and 3.2–3.5 cm diameter, pink to dark purple, tube paler inside, limb entire to shallowly 10-lobed, reddish pink, completely glabrous outside; stamens included, equal, 32–35 mm long, filament bases dilated and hairy, anthers 6–7 mm long; pistil included, disk cupular, 5-lobed, almost enclosing ovary, ovary glabrous, 2-celled, style 33–40 mm long. Fruits not seen.

Thailand.— SOUTHWESTERN: Phetchaburi [Kaeng Krachan National Park, 1 Sept. 2002, Traiperm 31 (A, BCU, BKF, K, KKU), Kaeng Krachan National Park, along road from Ban Krang camp to Khao Phanoen Thung ranger substation, 12 Aug. 2002, Middleton et al. 998 (A, BISH, BKF)].

Distribution.— Known only from the south-western region of Thailand in Phetchaburi province, collected twice in Kaeng Krachan National Park.

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