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THAI FOREST BULLETIN (BOTANY) 36

Igersheim & Ueachirakan 900914–1/5 (BKF); Ton Ouat FaWaterfall, 19 Oct. 2004, Kesonbua 110 (KKU)]; Narathiwat [no locality, 12 Jan. 1986, Niyomdham 1131 (AAU, BKF, P)].

Distribution.— India, Cambodia, Peninsular Malaysia, Singapore, Borneo.

Ecology.— Common in lowland evergreen and secondary forests, also along trails in open areas, forest edges, by streams and in coastal hill forest; altitude to 450 m. Flowering and fruiting periods: September to June.

Vernacular.— Chan thana bai lek (´µÄÁ¨È) (Narathiwat); khem pa (ÁȤiµ) (Narathiwat); ya-rong (¥µÃ¦) (Malay-Narathiwat).

Notes.— Tarenna wallichii is very variable as regards the shape of the leaves, which are oblong to lanceolate with inconspicuous lateral veins. The species is similar to T. quocensis, but can be distinguished by its broadly triangular calyx lobes and its smaller fruits with short persistent calyces. Its distibution, moreover, is restricted to the south of Thailand.

ACKNOWLEDGMENTS

We would like to thank the Faculty of Science, Ubon Ratchathani University and the Department of Biology, Faculty of Science, Khon Kaen University for supporting this study. Thanks to Dr. D.A. Simpson for his valuable advice and to anonymous reviewers for their numerous very helpful comments on the manuscript. We also thank the directors, curators and staff ofAAU, BK, BKF, BM, CMU, K, KKU, L, P, PSU, QBG,TCD and Herbarium of Biology Department, Chiang Mai University, for making specimens available for this study.

REFERENCES

Andreasen, K. & Bremer, B. (1996). Phylogeny of the subfamily Ixoroideae (Rubiaceae). Opera Botanica Belgica 7: 119–138.

Bremekamp, C.E.B. (1966). Remarks on the position, the delimitation and subdivision of the Rubiaceae. Acta Botanica Neerlandica 15: 1–33.

Bridson, D.M. (1979). Studies in Tarenna sensu lato (Rubiaceae subfamily Cinchonoideae) for part 2 of ‘Flora of Tropical East Africa: Rubiaceae’. Kew Bulletin 34 (2): 377– 402.

Craib, W.G. (1915). Contributions to the Flora of Siam. Additamentum 8. Bulletin of Miscellaneous Information, Kew 10: 425–437.

______. (1932). Florae Siamensis Enumeratio. A List of the Plants Known from Siam with Note of Their Occurrence.Vol. 2, part 1: Caprifoliaceae & Rubiaceae (in part). Siam Society, Bangkok.

De Block, P. (1997). Biosystematic Studies in the Tribe Pavetteae (Rubiaceae-Ixoroideae). PhD. Thesis, Department of Biology, The University of Antwerp, Belgium.

De Block, P., Degreef, J. & Robbrecht, E. (2001). Reinstatement of theAfro-Malagasy genus

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