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United States Department of Agriculture - page 14 / 36





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Description of the biology of the nonmodified recipient organism should include taxonomy, genetics, pollination, evidence of reported weediness (e.g., noting whether the crop or sexually compatible species is listed in the relevant publications of the Weed Society of America), and discussion of sexual compatibility with wild and weedy free-living relatives in natural crosses or crosses with human intervention. The applicant should provide the source of recipient (cultivar name or accession number) and the weed status of its sexually compatible relatives.

The applicant should explicitly identify the lines to be considered in the petition and the cultivars from which they are derived. If there are multiple lines, each line must be given a unique identifier and listed in the application. For virus-resistant plants, applicants should provide in an additional section the following information on the nature of the virus that provided the sequences encoding the resistance phenotype:

i) the taxonomic name of the virus including family, genus, and strain designation including any synonyms;

  • ii)

    the type of nucleic acid contained in the virus;

  • iii)

    whether the infection is systemic or tissue specific;

  • iv)

    whether the virus is associated with any satellite or

helper viruses; v) the natural host range of the virus; vi) how the virus is transmitted; vii) if transmitted by a vector, the identity of the vector including mode of transmission (e.g., persistent or nonpersistent); viii) whether any synergistic or transcapsidation interactions with other viruses under field situations have been reported in the literature, and situations have been reported in the literature; and ix) the location and the name of the host from which the plant the virus was originally isolated.

The above information can be provided in a table format (see Table 1). This information can be supplemented by listing references that report the host range, insect vectors, etc., for the virus.

Vegetative propagation is not a common mechanism by which cotton reproduces. Movement of genetic material by pollen is possible only to those plants with the proper chromosomal type, in this instance only to those allotetraploids with AADD genomes. In the United States this group would include only the cultivated species G. hirsutum, G. barbadense, and the wild species G. tomentosum. G. thurberi, the native diploid from Arizona with a DD genome, is not a suitable recipient. Movement to G. hirsutum and G. barbadense is possible if suitable insect pollinators are present and if there is a short distance from transgenic plants to recipient plants. Physical barriers, intermediate pollinator-attractive plants, and other temporal or biological impediments would reduce the potential for pollen movement.

Movement of genetic material to G. tomentosum is less well documented. The plants are chromosomally compatible with G. hirsutum, but there is some doubt as to the possibility for pollination. The flowers of G. tomentosum seem to be pollinated by moths, not bees, and the flowers are receptive at night, not in the day. Both these factors would seem to minimize the possibility of cross-pollination. However, Fryxell (1979) reports that G. tomentosum may be losing its genetic identity from introgression hybridization of cultivated cottons by unknown means.

  • G.

    Characteristics of Nontransformed Cultivar

  • G.

    hirsutum L. cv. “Stoneville 825” is the cultivar that we

genetically transformed. This cultivar is widely grown in the United States and was specially developed for introduction in the Mississippi Delta region. American Star Inc. has received a U.S. patent on the specific herbicide tolerant gene that has been transformed into this cultivar, and the transformed cultivar has received additional protection under the Plant Variety Protection Act of 1970. American Star Inc. intends to introduce the new traits into other cotton cultivars by traditional breeding techniques. The cultivar that has been transformed to be herbicide tolerant is called “Banjaran.”

SAMPLE PETITIONHerbicide-Tolerant Plants

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